Salinity Responsiveness in Finger Millet Analysis

Salinity Responsiveness in Finger Millet Analysis Introduction Salinity represents a strong limitation for agricultural production worldwide, especially in arid and semi-arid and restricts efficient utilization of available land resources. It is estimated that about 7% of world agricultural land that nearly one half of the total area of irrigated land could be adversely affected by salinization (Kosova 2013). Most of the cereal crops are sensitive to salinity and have limited amount of genetic variation for salinity tolerance in their germplasm. Hence genetic improvement of crops for their tolerance against salinity will be helpful in achieving targeted food production to meet the demands of growing population. Conventional plant breeding approaches have resulted in limited success in developing salt tolerant crop varieties due to multigenic nature of salt tolerance mechanisms and presence of low genetic variation in major crops. Another problem associated with conventional breeding is that if the gene is present in a wild relative of the crop, there is difficulty in transferring it to the domesticated cultivar, due to reproductive barriers and linkage drag. Recently, substantial progress in elucidation of salt tolerance mechanisms, especially salt ion signaling and transport, has been achieved due to utilization of modern genetic approaches and high-throughput methods of functional genomics. Genetic engineering has been demonstrated to be successful in developing salt tolerant crop plants (Zhang et al. 2001; Su and Wu 2004; Zhang et al. 2001). Genetic engineering strategies targeting various metabolic pathways viz., accumulation of osmolytes, antioxidant enzymes and up regulation of genes involved in stress responses like ion transporters, ion channels, transcriptional factors and various signaling pathway components have resulted in production of genetically modified crop plants exhibiting improved level of salinity tolerance (Turan et al. 2012). Identifying novel genes, analyzing their expression patterns in response to salt stress and determination of their potential functions in salt stress adaptation will provide the basis for effective genetic engineering strategies to enhance tolerance against salt stress (Cushman and Bohnert 2000). Responses against salinity stress involve many molecular processes such as ion homeostasis (membrane proteins involved in ionic transport), osmotic adjustment and water regime regulation (osmolytes) and scavenging of toxic compounds (Munns and Tester 2008). During recent years, considerable attention has been given towards elucidating the molecular basis of salt tolerance in crop plants. Several important pathways involved in salinity tolerance have been identified in model plants like Arabidopsis and rice (Zhu 2003; Walia et al. 2005; Cotsaftis et al. 2011). It is hypothesized that exploitation of halophytes or distantly related crops or wild progenitors of cereal food crops exhibiting superior levels of salinity tolerance may lead to identification of novel metabolic pathways/mechanisms/genes involved in modulating salinity stress tolerance in crop plants. Several research groups are working on understanding mechanisms of salinity tolerance in Pennisetum glaucum (Mishra et al. 2007), Avecinnia marina (Mehta et al. 2005), Porteresia coarctata (Garg et al. 2014) with a view to identify novel genes for genetic engineering of salinity tolerance in crop plants. But much more concerted efforts are needed to identify and exploit diverse crop species exhibiting superior level of salinity tolerance which will help in identifying novel genes associated with salinity tolerance. Finger millet (Eleusine coracanaL.) is an important minor cereal crop widely grown in Africa and Asia, known for its high degree of tolerance against drought, salinity and blast disease (Shailaja and Thirumeni 2007; Agarwal et al. 2011). Investigating the mechanisms and pathways involved in salt-tolerance of finger millet could facilitate better understanding of the molecular basis of salt tolerance and therefore enable the effective use of genetic and genomic approaches to improve salt tolerance in major cultivated crops. Although a wide range of significant physiological mechanisms and genetic adaptations to salinity stress has been observed, the underlying mechanisms of salt-tolerance in plants are still poorly understood. The best possible approach to explore tolerance mechanisms is to compare the components involved in stress response in tolerant as compared to sensitive plants. The other alternative to overcome this limitation would be to pick up some selected conserved genes w hich may be used to perform limited transcriptome analysis among the diverse genotypes. With this background, we planned to understand the physiological and molecular basis of salinity responsiveness in finger millet in comparison to the major cereal food crop, rice. Comparative physiological studies were conducted with a view to prove the superiority of finger millet genotypes over rice in terms of salinity tolerance. Two contrasting finger millet genotypes were used for physiological studies and expression analysis of already identified salinity responsive genes was done. This is the first study conducted to compare molecular basis of salinity tolerance in finger millet with rice. Material and Method Genetic Materials Used Seeds of two contrasting genotypes of rice (Oryza sativa) {FL478 (tolerant), White Ponni (Susceptible)} and finger millet (Eleusine coracona) {Trichy 1 (tolerant), CO12 (Susceptible)} in terms of salinity tolerance were evaluated for their responses against salinity stress under greenhouse conditions. Nucleus seeds of rice genotypes were obtained from Paddy Breeding Station, Tamil Nadu Agricultural University, Coimbatore, India and finger millet genotypes were obtained from Millet Breeding Station of Tamil Nadu Agricultural University, Coimbatore, India. Effect of salinity stress during germination Contrasting genotypes of rice {FL478 (tolerant), White Ponni (Susceptible)} and finger millet {Trichy 1 (tolerant), CO12 (Susceptible)} genotypes were assessed for their ability to germinate under salinity stress. Twenty seeds of both rice and finger millet genotypes were allowed for germination under different concentrations of NaCl solutions (0 mM, 50 mM, 100 mM, 200 mM NaCl solution) in petri-dishes with adequate replications. Germination percentage was calculated based on the number of seeds successfully germinated and vigor index was calculated based on the shoot length and root length on 10th day of germination. Effect of salinity stress during vegetative stage Imposition of salinity stress Contrasting genotypes of rice and finger millet genotypes (three seedlings per pot) were grown in perforated pots of 15 cm diameter and 20 cm height (having 3–5 mm holes on the side walls and bottom) filled with 2 kg of field soil mixed with required amount of fertilizer [1.25 g of (NH4)2SO4, 0.08 g Muriate of potash (KCl), and 0.08 g single superphosphate (SSP)]. Three pots were placed inside a large tray containing irrigation water and grown up to 20 days under greenhouse conditions. Plants were grown during June–August when air temperature ranged from 26 to 34  °C during the day and from 20 to 27  °C during the night and relative humidity ranged from 60 to 80 %. Salinity stress was imposed on 21st day when plant has reached to 5 leaf stage by adding desired concentrations of NaCl viz. 150 mM and 300 mM along with suitable control pots irrigated with normal water. Progression of salinity stress was monitored by periodically measuring the electrical conductivity (E C) of soil (from pot) and water (collected from tray) samples collected from both control and salinity stressed trays. Physiological and biochemical responses of contrasting rice and finger millet genotypes under salinity stress Contrasting genotypes of rice viz., FL478 (tolerant) and White Ponni (susceptible) and finger millet viz., CO 12 (susceptible) and Trichy 1 (tolerant) were evaluated for their physiological and biochemical responses viz., osmotic tolerance ability, salt accumulation pattern and sugar accumulation pattern during salinity stress. Measurement of Osmotic tolerance ability For assessing the osmotic tolerance ability of contrasting rice and finger millet genotypes, freshly emerged leaf (5-6cm) was marked and increase in leaf length was measured at every 24hrs interval during the initial 6 days of salinity stress along with control plants. Terminal leaf elongation rate per day (24 h) was calculated based on the observations recorded. Salt accumulation pattern Salt (Na+ and K+) uptake, transport and accumulation pattern of contrasting rice and finger millet genotypes was assessed by determining the (Na+ and K+) contents in shoots and top 3 leaves collected under normal and salinity stress conditions. Tissue samples collected at 21 DAS (days after stress) were washed with de-ionized water, dried in a hot air oven (70  °C) and then ground into fine powder. Ground samples were digested with triple acid mixture (sulfuric acid, perchloric acid and nitric acid in the ratio 9:2:1 v/v). Na+ and K+) concentrations in the triple acid digested extract were estimated using Flame Photometer (Elico, CL378). Determination of total soluble sugar content Total soluble sugar (TSS) content in the top three leaves of control and salinity stressed plants (21 days after stress) of contrasting rice and finger millet genotypes was determined using anthrone reagent method (Yemm and Willis 1954). Fresh leaf sample (100 mg) was ground in liquid nitrogen and pigments were removed using acetone extraction. TSSs were extracted in 80 % ethanol and were estimated by the anthrone reagent method using glucose as the standard. Other physiological responses of contrasting finger millet genotypes to salinity stress Gas exchange parameters were recorded in the third leaf (from top) of control and salinity stressed plants of rice and finger millet genotypes between 1000 hours and 1200 noon at 11 DAS (days after stress) using LI-COR 6400-XT photosynthesis system (LI-COR Biosciences, Nebraska, USA). The instrument was set with the following conditions: photo-synthetically active radiation 1,500  µmol of photon m−2s−1; ambient levels of CO2 and temperature; leaf area 3 cm2 and flow rate of 500  µmol s−1. RNA isolation, Northern blotting and hybridization Expression analysis of already reported salinity responsive candidate genes in response to salinity stress in the leaves of contrasting rice and finger millet genotypes were studied by northern blotting. Top 3 leaves of both rice and finger millet genotypes were collected and frozen immediately in liquid nitrogen from both control and stressed plant (300mM NaCl) when susceptible rice variety viz. White Ponni has shown salinity symptoms i.e., 11 days after salinity stress. Total RNA was isolated from stressed and control leaf samples using One Step RNA Reagent (Biobasic Inc., Canada) as per manufacturer’s protocol. The integrity of RNA was assessed by formaldehyde agarose gel electrophoresis. Total RNA was quantified using Nanodrop ND-1000 spectrophotometer (Thermo Fisher Scientific, Wilmington, DE, USA). 20ug of RNA mixed with RNA loading dye (1:1) was denatured at 75à ¢Ã‚ Ã‚ °C for 10mins and separated on denaturing agarose gel as described by Streit et al. (2008). The gel was stained with ethidium bromide and photographed. Gel was processed and RNAs were transferred to positively charged nylon membrane (Pal Corporation) using 20XSSC buffer. After capillary transfer to the membrane, RNAs were fixed by exposing the membrane to UV cross linker (Hoeffer, Piscataway). DNA fragment of candidate genes to be used as probe were isolated from rice cloned in pTZ57R TA cloning vector and confirmed by sequencing. Double-stranded probes were radioactively labelled with (ÃŽ ±-32P) dCTP using DecaLabel DNA Labeling kit (Fermentas) and probes were purified using Sephadex G-50 spin column (GE Healthcare). Radiolabelled probes were denatured on boiling water bath snap cooled on ice and used for hybridization as described by Streit et al. (2008). RNA blots were pre-hybridized in ULTRAhyb ® at 45à ¢Ã‚ Ã‚ °C for 4–8 h. The blots were hybridized with 32P-labelled denatured probes at 45à ¢Ã‚ Ã‚ °Cfor 20 h in the same but fresh buffer. The blots were initially w ashed at room temperature with 2XSSC and 0.1% SDS followed by twice wash with 1XSSC and 0.1%SDS at 45à ¢Ã‚ Ã‚ °C for 20 min each. The blots were initially washed at room temperature with 2XSSC and 0.1% SDS for 30 min and then washed with different stringencies for different probes to decrease background. Hybridized membrane were dried on blotting paper and exposed to Kodak XAE-5 film with cassette having Kodak intensifying screen for 1–6 d. The resulting radiograms were scanned in an LKB 2201 densitometric scanner. Results Effect of salinity stress on rice and finger millet genotypes during germination stage Screening of contrasting genotypes of both rice and finger millet against salinity stress at germination stage revealed the superiority of finger millet over rice in terms of salinity tolerance at germination stage. At lower concentration of salinity stress (i.e 50mM NaCl) the susceptible genotypes of both finger millet (CO12) and rice (White ponni) has shown better germination percentage and vigor index as compared to tolerant genotypes. Tolerant rice genotype FL478 was found to possess better germination percentage (35 ±2.9%) and vigor index (128.2 ±10.6) in comparison to susceptible White Ponni where germination percent and vigor index was found to be 16.7 ±1.7% and 70.9 ±7.1 respectively. Both finger millet genotypes i.e. CO12 and Trichy1 has shown almost similar germination percent and vigor index at 100mM of NaCl stress. Both rice genotypes (viz. FL476 and White Ponni) did not show any germination beyond 100 mM NaCl stress (Table 1); whereas both susceptible (CO12) and tolerant (Trichy 1) finger millet genotypes were able to germinate even at 300 mM NaCl stress (Table 1). At 300 mM NaCl stress Trichy 1 has shown better germination percent (40.0 ±1.6) and vigor index (32.0 ±1.3) as compared to CO 12 germination percent (24.4 ±0.9) and vigor index 24.4 ±0.9.

With reference to empirical examples, explore the everyday activities of different groups of migrant workers in (re)producing processes of ‘globalisation’

Globalisation is most commonly known as the process that combines international practices, falling into various strands consisting of â€Å"politics, economics and culture as well as mediated communications† as stated by Marsella and Ring (in Kofman and Youngs, 2003: 3). There are many definitions of globalization that are widely recognized in our present day, the most common, which I will be referring to in this essay, being that of greater economic integration by the increased unity of our world, also known as ‘Economic Globalisation'. Harrison (2007:35) refers to Economic Globalisation as â€Å"increases in trade, foreign investment, and migration† and according to the Global Education Program, within the past few years, the improvement of technologies and ‘reduction of barriers' has meant that the level of exchange between people and countries in terms of ‘goods, services, knowledge and cultures' is ever increasing at speedy rates. As migration falls into a factor of economic globalization, in this essay, I plan to explore how migration influences globalization and the impact it is having on the migrants themselves, also taking into account the social dimension of globalization in terms of how it affects the migrant's identities socially and culturally. Being one of the most passionately debated issues in politics today, Globalization is often split in outlook, with those ‘for' and those ‘against' the phenomena. The positive believers of Globalization argue it is the foundation for solving problems of high unemployment and poverty worldwide, whilst others believe it is rather the catalyst of these issues. In terms of the benefits of Globalisation on an international scale, Dinello and Squire (2005; xv) states â€Å"the proponents of globalization often point to its three-fold beneficial impact, with positive implications for equity: 1) stimulating trade and economic growth, 2) reducing poverty without rise in inequality and 3) contributing to economic and political stability†¦ and statistics provided strongly support these claims. For example, Uganda in 1990 had its poverty rates fall by around 40 per cent whilst its rate of school enrolment doubled in numbers (Dinello and Squire, 2005:xv). However, those who are â€Å"anti-Globalisation† equate the process as having negative impacts such as inequality by only creating economic growth in selected countries as well as increasing the rate of vulnerability in countries and people. Used as an example is China, which has had a remarkable success since entry into the ‘global economy†, however, this success has been accompanied by an â€Å"unparalleled rise in the country's within-country inequality† (Dinello and Squire, 2005:xvi) Migration, as stated by Marsella and Ring (in Adler and Gielen, 2008:11), refers to â€Å"the act or process by which people, especially as a group, move from one location†¦ to another† being a procedure that has been an ‘inherent part of human existence' from early centuries, it is now a central form of our ‘global flow of persons, goods, practices and ideas'. According to the International Organization of Migration (IOM) migration is currently viewed as being one of the most ‘defining global issues' in our present day, with around 192 million people living outside their place of birth in 2005, accounting to be roughly 3 percent of the worlds overall population. The annual growth rate of migration being around 2. 9 percent, however, it should be noted that this rate has stayed constant over the past 30 years (Van Hear, N. nd Nyberg-Sirensen, N, 2006). In continuation, an important factor that Marsella and Ring (2008) raise is the ideology that the movement of migration arises from what can be referred to as â€Å"push and pull† factors. Putting it into perspective, the pull factors of migration being the option of ‘new possibilities, ‘rewards' and ‘hope' as a majority of migrants leave their country of origin to increase their economic prospects and be reacquainted with family and friends. On the other hand, the push factors, which cause migrants to leave their home countries being that of â€Å"a sense of danger, discontentment and boredom† or avoiding dangers of persecution in their home countries (in Adler and Gielen, 2008:11). Van Hear, N. and Nyberg-Sirensen, N (2003: 51) state â€Å"The ratio of real income per head in the richest countries to the poorest rose from 10:1 in 1900 to 60:1 by 2000. Such disparities in living standards and the lack of development options in developing countries are at the root of much migration. † Western Europe, North America and Australia are more recognized as the preferred locations by all migrants ranging from the lower class, asylum seekers and skilled professionals. However, despite migrants personal beliefs of achieving better lives, historically, migratory workers are most typically known to have very low economic status due to their low pay, often living in ghettos and suffering from relative poverty due to migrants being â€Å"sources of cheap labour†, vulnerable to abuse, exploitation and mistreatment (Adler and Gielen, 2008:11). Frequently, workers from developing countries such as Bangladesh and Vietnam are lured to developed countries with the illusions of being provided with better economic status through higher wages and improved employment only to arrived and be subjected to wages below the minimum and inhuman working hours. Marsella and Ring (in Adler and Gielen, 2008:11), referred to this as ‘modern-day slavery', as often these migrant workers can not â€Å"escape, and if they try may be assaulted or killed (as)†¦ Many are illegal immigrants†¦ and have no one to turn to for assistance† The process of Migration, however, has an impact not only on those directly involved but also on their host country, the biggest benefit, being the transfer of skills and labour as countries can invite migrants to fill in the gaps in their labour markets. In addition, they introduce new cultures, increase workforce competition but can also have negative impacts such as increasing strains on the host's economy if claiming government benefits. Although Migration is a key element greatly influenced by Globalisation, it is found that â€Å"Discussions of Globalisation rarely consider international migration at all†¦ † (Stalker, 2000;1). In terms of how migration relates to Globalisation, Taran (1999) implies that this new age of Globalization is â€Å"now generating a new configuration of forces promoting migration† (Adler and Gielen 2008:13) and states seven reasons for current Globalisation influenced migration, some of which being â€Å"aspects of Globalisation such as unemployment and culture conflict†,† development-induced migration† and â€Å"large-scale corruption. This being highly portrayed in the fact that increases in â€Å"forced migration† is present as the number of â€Å"illegal immigrants showing up throughout the world and requesting asylum† is on the increase. An example being that of hundreds of illegal migrants that had to be declined entry into Australia via Indonesia typifies (Adler and Gielen 2008:15). There are a lot of misconceptions about the current trends in migration and development, Van Hear and Nyberg-Sirensen (2003: 51) argue that â€Å"the popular conception that the poor are migrating from the (Global) South to the (Global) North is unfounded†. In fact it is shown that â€Å"most migration is among developing countries rather then from the developing world to the developed†. An important fact they raise is that of the lack financial resources that ‘the poorest of the poor† have, which is ‘1. 2 billion people living on less than US$1 a day' as well as adequate network connections, they simply can not afford the cost of ‘Inter-continental migration'. The ideology of migrant workers being predominantly in the unskilled work sector is also debatable. A good example of being Switzerland, which, due to a raise in economic expansion and the need for workers to fill the labour shortages, signed foreign labour recruitment agreements and now is classed as having one of the highest immigration rates on the continent. Similar to most European countries, Switzerland has and continues to take advantage of the available labour in other countries to ensure economic growth. In Accordance with the 2000 census, over 22% of its total population were foreign born. In continuation, although the beliefs are that migrants tend to have low status jobs, the Organisation for Economic Co-operation and Development (OECD) state that in 2001, the highest number of foreign workers were â€Å"in the areas of commerce, banking and insurance† in Switzerland, which was a figure just over 24%, next was metalwork and machinery (15. 8%) (OECD, 2004; 36). The statistics suggest traditional views of migrants being low-end workers may be rather a narrow viewpoint, especially in terms of countries within the European Union. In saying this however, although the lack of available statistics makes it difficult to assess the skill levels of foreign workers in Switzerland, statistics of available data does suggest that a high majority of foreign workers are unskilled in relation to Swiss Nationals. A high 36% percent of Foreign nationals leaving school after compulsory education, in comparison to only 18% for Swiss nationals (OECD, 2004). Migration has positive effects on two levels,Van Hear, N. nd Nyberg-Sirensen, N describe this as the ‘family level' and the ‘community level'. The family level being the fact that migration can help people gain access to better housing increased household earnings and advanced healthcare and educational opportunities. Van Hear, N. and Nyberg-Sirensen, N 2003: 52) argue the â€Å"positive effects may spread to the wider community and society, preventing the decline of rural communities or collapse of national economies†. In terms of the community level, â€Å"migrants' hometown associations (HTAs) may serve as platforms resulting in significant development, such as improvements in local health, education, sanitation, and infrastructure conditions, benefiting migrant and non-migrant households. â€Å". Van Hear, N. and Nyberg-Sirensen, N argue however that a major drawback of migration is that the negative impact faced by the ‘sending community' if a majority of their skilled labour force is sent abroad, this devaluing the countries labour market. Nevertheless, through remittances this disadvantage can be overlooked, as it allows surpluses to be sent back to home families. Studies have shown that remittances generally contribute to a raise in trade levels, income distribution and economic growth, Kavita Datta et al (2007: 46) refers to the â€Å"growing focus on financial remittances as potential drivers of development† where in a recent report made by the World Bank, â€Å"officially recorded remittances to the developing world in 2005 were US$167 billion, a dramatic increase from US$31. 2 billion in 1990† Therefore making remittance â€Å"the second-largest capital ? w – behind foreign direct investment†. Remittances have a great impact on those in developing countries as they provide immediate financial support. Manuel Orozco (2002) argues that the function of recent family remittances is a vital sign of Globalisation in Central America and the Caribbean. Manuel (2002) states â€Å"family remittances are currently one of the most important forms of linkage among emigrants Latinos and Latin America† confirming that â€Å"Many Latin American countries find family remittances an important source of national income† (Manuel, 2002;46). Din (2006) stated â€Å"remittances continue to play an important part in the link between ‘prosperous' British Pakistanis and those relatives who still depend on remittances†. It is especially important for many households, as it is a direct form of financial resources and therefore has a greater impact then other resource flows (N. and Nyberg-Sirensen, N 2003: 53). Datta et al (2007; 53) refers to a care worker from Jamaica who regularly sent money back home and a man who has taken on the role of a ‘transnational father' sending home not only financial remittances for his children but also various needed resources. On the other hand, As beneficial as remittances are they are also selective and do not benefit the wider community, tending to be most advantageous to those from better-off households who have the initial capital to send that family member abroad (N. and Nyberg-Sirensen, N, 2003). Din (2006; 25) points out that it mainly only benefits the immediate family and in relation to Pakistanis â€Å"despite the increase in financial resources for some lower caste families, they still remained in the same caste†, meaning they still faced the same judgments. The cultural impact migrants have on Globalisation is highly evident in Britain today, where we have a high range of various ethnic minority and exceedingly multi-cultural communities, creating a versatile and vibrant society, full of a vast range of cultures and identities. Often migrant's practices, customs and branches of their original culture are brought over and recreated in their host countries, an empirical example being that of Chinatown in London's West End. Although there are many advantages of cultural migrant impacts, feelings of hatred can arise and prompt social conflicts. Moses (2008:176) argues that while â€Å"many people are willing to embrace the economic and political gains from globalization, there is a persistent fear that globalization undermines national cultures and identities. † A multicultural society may be seen as quite threatening to those native-born citizens of the host countries. Engels (in Ikhlaq Din, 2006:29) states â€Å"migration has always been a controversial issue both socially and politically. During periods of mass migration to the UK white and non-white migrants faced hostility when looking for employment and housing†. However, In Margaret Brearly's (2007) article â€Å"THE ANGLICAN CHURCH, JEWS AND BRITISH MULTICULTURALISM† she states the statistics collected from a mori poll for the BBC in August 2005, soon after the London July bombings showed that although 32% of the population thought that multiculturalism â€Å"threatens the British way of life†, 62% believed that â€Å"multiculturalism makes Britain a better place to live. Its can be argued that Migration causes the involved individuals to lose their own culture in some cases. For example, Margaret Brearly's (2007) refers to ‘Other commentators' that have argued that multicultulist policies that have failed to â€Å"to promote formal learning of English, prevent integration† which has caused â€Å"inter-ethnic tensions and ghettoization into separate enclaves with high unemployment and social alienation. (75% of Pakistani and Bangladeshi families are classified as living in poverty, while 35% of Muslim children live in ‘workless' homes)†¦ In this, following this concept, through the movement of increased migration and globalization as a whole, there would be greater harmonization, which could evidentially lead to one overall shared culture worldwide. Datta et al (2006; 48) state it is evident that the process of migration is becoming a highly influential factor for the functioning of global cities such as London, â€Å"arguably creating a ‘migrant division of labour' (May et al. , 2006)†. Work Permits (UK) has facilitated the application process so for example, foreign students studying the United Kingdom have the ability to apply for a work permit immediately after they graduate, whereas previously they were required to return to their home country before application (Organisation for Economic Co-operation and Development; 2004) Datta et al (2007) quotes Ratha (2003) in stating â€Å"there has also been a major shift in recent years in the way in which Northern governments view migration†. For a start, a number of governments and IFIs now agree that migration contributes to economic growth both nationally and globally†. In â€Å"International migration† by Jonathon Moses (2006; 159) he refers to two groups of migrant home countries, those with a â€Å"relatively small, but poorly paid workforce† and those with a â€Å"limitless supply of cheap unskilled labour†. The latter group being that of countries such as Bangladesh and India, which have an infinite amount of cheap ‘unskilled labour'. In these countries, it is expectant to find government incentives and ‘targeted government institutions' to encourage ‘migration, remittances and repatriation'. Moses (2006) goes on to explain that in the countries without excess surplus of labour, emigration can reduce the nations already limited supply of labour, not all negative however, â€Å"this increased scarcest brings with it greater influence and a better price†. Therefore internationally it can be argued that migration has lead to improvements in both the Global South and North as Harrison (2007) argues that poverty in the South falls for two reasons, firstly â€Å"the migration of capital to poor countries raises wages in poor countries, and the migration of unskilled labor from poor to rich nations raises the income of both the migrants and those workers who remain behind† (Harrison, 2007; 112). However it is also important to note that many developing countries face what Moses (2006) refers to as â€Å"brain drain† which was first used to refer to the Indian Economy. During the 1970s to 1980s it was found that the ‘entire graduating classes' from elite Indian institutes of technology emigrated, many achieving financial success abroad (Moses; 2006). Africa being the worst disadvantaged, as it suffered from a shortage of labour, a substantially high number of doctors trained in Ghana during the 1980s left the country, according to the UNDP's 1992 Human Development report, the figure was at least 60 per cent. The shortage of labour therefore, leading to a stunt in economic development (Moses 2006). However, Moses (2006; 174) suggested that in the long run these home countries could benefit from the ‘brain-drain' thesis as first generation â€Å"brain-drain migrants have managed to build technology bridges that span the divide separating developing and developed worlds. † Once the bridges are created these individuals can then return back to their countries and capitalizes on investments that have been made in education and human ‘capital development' (Moses 2006; 174). To conclude the main influence migrants have on Globalisation is through remittances, by supplying their families back home with financial support they aid in lowering the rate of poverty. In addition, if remittance funding is used for capital generation it aids in creating an increase in the home countries economic growth. However, due to the nature of remittances being selective, it is not beneficial to all and could be argued promotes further inequality, especially within developing countries. In addition, the low working conditions and personal sacrifices these workers make to be able to remit needs to be put into consideration. What needs to promoted is the economic development of the Global South, which would decrease the incentives to migrate as, in accordance with Stalker (2000;10) â€Å"In theory Globalization should eventually make countries economically more equivalent so people should not need to move around the world searching for work† . More development policies are needed and perhaps the advancement in Globalisation is needed, as the exposing of developing economies to the Global world, through competition, will create efficiency and productivity. The detrimental effects of unsuccessful migration also needs to be taken into account, in cases where migrants are not contributive to taxes and drain on government revenue and in terms of culture where native-born citizens feel threatened. All in all, i feel for migration to have an influential impact on Globalisation trade and foreign investment must also be taken into account, as migration is only one factor. Secondly for a positive impact, migration must be controlled and efficiently regulated.

The Ugly Secret of Argumentative Essay Samples for Teachers

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Imperialism and China - 2269 Words

Imperialism’s effects on China has always been controversial and debatable. Andrew Nathan’s article on Imperialism’s Effects on China explains imperialism’s positive contributions to modern day China and defends the theory that China’s difficulties are caused by internal factors rather than external. On the other hand, Joseph Esherick’s Harvard on China: The Apologetics of Imperialism argues that China’s economic and social disruptions are caused by imperialism. However, when one examines the evidence chosen and rhetoric of both sides, one may realize the faults in Nathan’s essay which are not shown in Esherick’s. Esherick, as opposed to Nathan, delivers his arguments and defends his stance in a well-rounded manner along with wise usage of†¦show more content†¦However, he does agree with Nathan on one positive effect of imperialism: â€Å"the very struggle to eliminate the economic, political, social, and psycho logical vestiges of imperialism produced the basis of sustained, self-reliant economic and political growth.† A thorough analysis of the weaknesses and strengths of each essay is necessary to prove that one side is more convincing. Nathan has two main strengths, the first is laying out the historical background. He mentions the six institutions established by the foreign nations – the treaty ports, spheres of influence, other restrictions on Chinese sovereignty, the financial drain, the missionary invasion. Under each topic, he acknowledges the negative impacts they had on China; for instance, â€Å"the atmosphere of the treaty ports were strongly racist†¦Ã¢â‚¬  â€Å"the treaties involved China in financial obligations to foreigners that were crippling to government finance.† With this evidence, the readers gain a brief understanding of the historical background. His second strength is clarity. He responds to and outlines the propositions made by his opp onents. He begins by responding to economic arguments on mass pauperization, then to the assumption of stifling economic growth in the late Qing dynasty, and the statement that imperialism causedShow MoreRelatedImperialism in India and China1234 Words   |  5 PagesBritish imperialism in China and India brought very different responses, in part because of the nature of imperialism in each place. While both regions were greatly influenced by the British, in India the country was placed under the direct rule of the Queen. 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